Applications now open. This 4 year PhD project is part of a competition funded by EASTBIO BBSRC Doctoral Training Partnership (DTP). Detailed guidance on the application process, and the EASTBIO Application and Reference Forms can be found at How to Apply | Biology. Please send your completed EASTBIO Application Form along with a copy of your academic transcripts (as a pdf) to student-admin@igc.ed.ac.uk. You should also contact your referees and ask them to submit their references (on the EASTBIO reference form template) to student-admin@igc.ed.ac.uk by the application deadline of noon on 15th December 2025.This opportunity is open to UK and international students and provides funding covering stipend and UK level tuition fees. The University of Edinburgh covers the difference between home and international fees meaning that the EASTBIO DTP offers fully-funded studentships to all appointees. There is a cap on the number of international students the DTP recruits. It is therefore important for us to know from the outset which fees status category applicants will fall under when applying to our University. Please refer to UKRI website and Annex B of the UKRI Training Grant Terms and Conditions for full eligibility criteria. Single cell proteomics to detect emergent cell states associated with ageing Supervisors: Professor Simon Wilkinson and Dr Alex von Kriegsheim Investigating the mutational consequences of transcription-damage interactions through computational modelling Supervisors: Professor Martin Taylor and Michael Nicholson (School of Mathematics) Dissecting the effects of gene body methylation on gene expression in mammals Supervisors: Dr Duncan Sproul and Professor Greg Kudla Using disease-causing and population variants to dissect ATRX protein function Supervisors: Dr Rebekah Tillotson and Dr Atlanta Cook Molecular Mechanisms Linking Nucleolar Function to Hypoxia, Inflammation and Aging Supervisor: Professor Lesley Stark Dissecting CIC::DUX4 oncogenic circuitry through single-cell perturbation profiling and network inference Supervisor: Dr Nezha Benabdallah This article was published on 2025-11-18