One-off opportunities and Edinburgh Doctoral College Scholarship Metabolic rewiring of Hepatocellular Carcinoma Supervisors: Professor Alex von Kriegsheim, Dr Rachel Guest, Dr Luke Boulter and Dr Stephen Chapman Investigating the niches of stem and progenitor cells in arthritis Supervisor: Professor Cosimo De Bari Dissecting the Spatial Origins of Biliary Intraepithelial Neoplasms Supervisors: Dr Luke Boulter and Prof Tim Kendall Edinburgh Cancer Research Funded Studentships Applications now closed for non-clinical studentships CRUK Brain Excellence Fellowships Applications Now closed MRC 3.5-year International Visiting PhD projects: Human Genetics, Genomics and Disease Collaborative PhD Projects between MRC Human Genetics Unit (University of Edinburgh) and Max Delbrück Center (Berlin, Germany). Applications Now CLOSED Edinburgh Doctoral College Scholarship Applications now CLOSED for 2025 intake Precision Medicine PhD Projects Next round of applications announced late Autumn 2025 This article was published on 2025-03-05