TRAM PhD Projects

A list of available PhD projects offered as part of the TRAM MB-PhD Programme are displayed below

PhD Project TitleSupervisor and Location
Investigating the transcription factor FoxO1 in synovial fibroblasts

Dr Fraser Collins: Institute of Genetics and Cancer, University of Edinburgh

Professor Cosimo De Bari: Institute of Genetics and Cancer, University of Edinburgh

Developmental programming of salivary gland immune cells in Sjögren’s Syndrome

Dr Elaine Emmerson: Institute for Regeneration & Repair, Centre for Regenerative Medicine, University of Edinburgh  

Dr Rebecca Gentek: Institute for Regeneration & Repair, Centre for Reproductive Health/Centre for Inflammation Research, University of Edinburgh

Methotrexate Osteopathy: Epidemiological and Mechanistic Insights into Drug-Associated Bone Fragility

Dr Barbara Hauser: Institute of Genetics and Cancer, Univeristy of Edinburgh

Dr Athina Spiliopoulou: Usher Institute, University of Edinburgh

Dr Erika Kague: Institute of Genetics and Cancer, University of Edinburgh

Identifying endotypes in monogenic osteoporosis using a multifaceted approach

Dr Erika Kague: Institute of Genetics and Cancer, University of Edinburgh

Dr Athina Spiliopoulou: Usher Institute, University of Edinburgh

Professor Stuart Ralston: Institute of Genetics and Cancer, University of Edinburgh

Professor Outi Makitie: Karolinska Institute and University of Helsinki

Investigating the role of Enpp1 deficiency in osteoporosis.

Dr Vicky MacRae, The Roslin Institute, University of Edinburgh

Professor Patrick Hadoke, Centre for Cardiovascular Science, University of Edinburgh

Dissecting the role of COPA mutations in fibro-inflammatory lung disease

Dr Karen Mackenzie, Institute for Regeneration and Repair, University of Edinburgh


Dr Nik Hirani, Institute for Regeneration and Repair, University of Edinburgh 


Professor Chris Lucas, Institute for Regeneration and Repair,  University of Edinburgh

Molecular regulation of stem and progenitor cell fate in the repair of skeletal joints.

Dr Anke Roelofs: Institute of Genetics and Cancer, University of Edinburgh

Professor Cosimo De Bari: Institute of Genetics and Cancer, University of Edinburgh

Project Summaries

Investigating the transcription factor FoxO1 in synovial fibroblasts (Collins & De Bari)

Developmental programming of salivary gland immune cells in Sjögren’s Syndrome (Emmerson & Gentek)

Methotrexate Osteopathy: Epidemiological and Mechanistic Insights into Drug-Associated Bone Fragility (Hauser, Spiliopolou & Kague))

Identifying endotypes in monogenic osteoporosis using a multifaceted approach (Kague, Spiliopolou, Ralston & Makitie)

Investigating the role of Enpp1 deficiency in osteoporosis. (MacRae & Hadoke)

Dissecting the role of COPA mutations in fibro-inflammatory lung disease (Mackenzie, Hirani and Lucas)

Molecular regulation of stem and progenitor cell fate in the repair of skeletal joints (Roelofs & De Bari)